There are, however, a number of what may be called “sub-maritime” plants, which are neither halophytes nor xerophytes, but seem rarely to be found more than a few miles from the sea. Well-known examples of these are the slender thistle (Carduus tenuiflorus), alexanders (Smyrnium olusatrum) (Pl. IV), fennel (Foeniculum vulgare) (Pl. XXXIV), and one of the mouse-ear chickweeds (Cerastium tetrandrum). Besides these, there is a much longer list of species which, though not confined to the coastal belt, are always far commoner there than farther inland. Storksbill (Erodium cicutarium) and buck’s-horn plantain (Plantago coronopus) (Pl. XXXVI) are familiar examples. None of these plants conform to any special type (there are a number of aquatic plants as well) and no explanation is at present forthcoming to account for their distribution (see here).
It is unavoidable in a book of this kind that it should contain a large number of lists of the plants usually found in the various habitats. These may well seem tedious to some readers who are unfamiliar with the appearance of many of the species mentioned, and I have therefore devoted Chapter 12 to giving brief popular descriptions of plants mainly confined to the coastal belt. I have also added notes on their distribution and their relative importance in the general vegetation. It should be clearly understood, however, that it is in no sense my object to provide a flora for identifying all the plants likely to be encountered along the coast, and I have not attempted to include those species which are equally common inland. A number of standard floras are mentioned in the Bibliography (see here), and one of these should be consulted when identifications are being made. At the same time, it is to be hoped that this chapter may prove useful for quick reference. The main object of this book, however, is to describe seaside vegetation as a whole, and to relate it as far as possible to the various habitats where it is found.
I have devoted a separate chapter to the vegetation found in each of the main habitats. The following summary will give some idea of particular portions of the coast with which we shall be concerned in each chapter:
1. SALT-MARSHES (Chapter 5)
The mud or sand of the inter-tidal zone on flat shores, which is sufficiently protected from violent wave-action to support vegetation, including areas only flooded by the highest tides.
2. BEACHES AND FORESHORES (Chapter 6)
A comparatively narrow zone along the tops of exposed beaches of any material, other than rocks, only reached by the highest tides.
3. SAND-DUNES. (Chapter 7)
All areas of sand, originating in material blown by the wind from the shore, from shifting open dunes just beginning to accumulate round individual plants, to old and mature deposits whose surface has become almost completely stabilised by a close cover of vegetation.
4. SHINGLE BEACHES (Chapter 8)
The beaches, bars or spits of water-worn pebbles, derived from rocks by wave-erosion and deposited by the sea on low-lying shores. These vary from highly unstable banks of stones, continually shifted by the waves, to the oldest beaches where the shingle is completely dormant and has long been isolated from any wave-action.
5. CLIFFS AND ROCKS (Chapter 9)
Rocky places above the high-tide mark or cliffs of any material, which are to some extent exposed to salt spray. Certain artificial habitats such as sea-walls, which are similarly exposed to spray, are included in this category.
6. CLIFF-TOPS (Chapter 10)
The strip of ground along the tops of cliffs, exposed to a certain amount of spray and often supporting some characteristic sub-maritime plants as well as maritime and inland species. When the tops are level this is usually quite narrow, but on steep slopes it may cover a large area and then differs from the cliff habitats in the previous chapter only in possessing more soil.
7. BRACKISH WATER (Chapter 11)
The swamps, ditches, lagoons, or estuaries of slow-moving rivers where the water remains brackish as a result of sea-water mixing with fresh water. These are often inhabited by a characteristic mixture of sub-maritime and inland aquatic plants.
It is hardly necessary to add that these habitats are by no means always sharply separated, but in many places overlap considerably. Sand-dunes, for instance, often grow up on the crests of shingle ridges, and salt-marshes develop easily behind the protection of shingle or sand-bars. As a result, the vegetation is often mixed up in a distinctly confusing manner—nowhere better seen than along the north Norfolk coast.
CHAPTER 2 THE PHYSIOGRAPHICAL BACKGROUND
A Résuméby J. A. STEERS
THIS BOOK is concerned with the ecology of the sea-coast and the seashore. The various types of ground that come under this broad title are subject not only to constant change but often to violent change. Even the hardest cliffs are comparatively unstable, and almost always subject to strong winds and storm-waves which may do much superficial damage, even if the actual rate at which the cliff retreats is, in terms of human life, extremely slow.
Great Britain has a remarkably long and intricate coastline and a long and varied geological history. Strata of nearly every period are well represented. These rocks, and the associated igneous rocks (also of very different ages) give the coast great interest and variety. We can observe the white and often perpendicular cliffs of the Chalk, the magnificent ranges of dark red cliffs of Old Red Sandstone in Caithness and Kincardine, the grey walls of Carboniferous Limestone with their flat grassy tops in west Pembrokeshire, the rapidly wasting cliffs of glacial deposits of north Norfolk and Holderness, the heavily glaciated cliffs of the whitish-grey Lewisian Gneiss, alternating with those of the brown and often spectacular Torridon Sandstone in north-western Scotland, and the granite cliffs of Land’s End and the Isles of Scilly. These are but a few examples from many; the point is that the rock type alone—quite apart from whether the beds are folded, broken, horizontal, or cut into diverse forms by marine or sub-aerial erosion—makes the coastline extremely interesting.
CLIFFS
In the preceding paragraphs the word cliff has been used but not defined. The Oxford English Dictionary gives the following meanings for the word:
1 A perpendicular or steep face of rock of considerable height, usually implying that the strata are broken and exposed in section; an escarpment.
2 (especially in modern use) A perpendicular face of rock on the seashore, or (less usually) overhanging a lake or river.
These definitions are descriptive, comprehensive, and do not presuppose any particular origin. Only too often it is assumed that because cliffs face the sea, they are wholly the result of marine erosion. The mere presence of plants on a cliff face is not, of course, evidence that no erosion is taking place; it may suggest that erosion is fairly slow, or that it may operate by large and infrequent slips. But a steep, grassy slope running down to near sea-level may be quite untouched by marine erosion. Some of the so-called Hog’s Back cliffs of north Devon are only affected by the sea in their lowest parts; their upper slopes have been produced in some other way (Pl. V).
Many cliffs are found at the back of a flat or platform where they are no longer washed by the waves. Along the Durham coast this is so, and still more along many miles of the coast of Scotland, especially in Galloway, Kintyre, Arran, Ayrshire, and many other districts. Sea-level, relative to the land, has altered since the platform was formed, and where the alteration is considerable, the old cliff may now be well away from the sea and its vegetation only indirectly influenced by it. The boulder-beach at the cliff-foot south of Duncansby Head in Caithness indicates a slight change of level, since the boulders are lichen-covered and are somewhat above the normal height which wave attack reaches. In the Gower peninsula, the limestone cliffs run right down to sea-level, often in unbroken slopes. But in many small inlets traces of raised beaches are found, and it follows that the outer cliffs, even if they do run down to and below sea-level, cannot wholly be the product of modern marine erosion. In parts of Scotland there are sometimes two or three old beaches found in the same place, often with a normal shingle flora. Between the mouth of the Findhorn and Burghead (Moray Firth) there is a great series of ridges, all well above the present sea-level. To-day the sea is cutting into them and making a shingle cliff fifteen or more feet high. Their natural vegetation closely resembles that of modern beaches, but has undergone changes as a result of its more stable position. In many other places around the Moray Firth, especially between Hilton of Cadboll and Tarbat Ness, and in various localities on the west coast, particularly on Islay, Colonsay, and Jura, magnificent expanses of shingle-flats and old cliffs are to be seen.
In other places the sea-level has risen relative to the land, so that the lower valleys are flooded, and what were cliffs are now submarine slopes. Much of the beauty of Pembrokeshire, Cornwall and Devon depends on the influx of the tides into long and intricate inlets like Milford Haven, Plymouth Sound, the Camel estuary and the mouth of the Dart. These were carved first by rivers and streams on the land, later drowned by a relative rise of sea-level. Only their outer cliffs have been modified by marine action; the sheltered inlets now often contain salt-marshes. The pulsation of the tides into their innermost recesses, and the intimate relation between land and sea vegetation create beauty.
In western Scotland there is another type of inlet, the sea-loch or fiord. Many of the narrow straits between islands or between the mainland and adjacent islands are similar. These lochs usually owe their present appearance to ice action which has scoured out, widened, and perhaps deepened and straightened pre-existing river valleys, which sometimes followed lines of weakness produced originally by faulting. The sides of these lochs and straits are cliffs in the sense of the dictionary definition, but are not the product of marine erosion. Up the fiords there are often salt-marshes, and at low water the expanse of golden, brown, red and varicoloured seaweeds, surrounded by the higher marsh plants, under a strong sun is unforgettable.
Cliff-form is usually closely related to structure. If the beds are more or less horizontal and thick only one may form the cliff, but more often two or more drop out. At Hunstanton the brown Carstone overlain by the Red Chalk and this again by the White Chalk makes a spectacular cliff, subject to rapid erosion because of the ease with which the sea eats into the soft Carstone, thus producing falls. Near Lyme Regis, and in Glamorgan, the nearly vertical cliffs are composed of thin beds of limestone and clays or shales arranged horizontally. Elsewhere, the beds may be steeply inclined seawards or landwards, folded or faulted, and the form of the cliff will depend much on the trend of the cliffs relative to that of the folds. The rocks of the Isle of Wight and the Isle of Purbeck are strongly folded in an east-west direction. The cliffs along their south coasts run with the beds which, seen from the sea, appear almost horizontal. But if one sails round Durlston Point to Poole Harbour, or between Ventnor and Ryde, the beds are seen on end, and the way in which the sea has cut into the softer ones is evident. Where folding is acute and intricate and the rocks are hard, the sculpture is often bizarre. A view from a boat close inshore between Boscastle and Hartland Point, or between Berwick and Cockburnspath, or around the west of Pembrokeshire reveals details of surprising interest.
Where all the rocks are soft and perhaps geologically young, changes go on at a quicker rate than where the rocks are harder and older. Changes are rapid between Flamborough Head (Pl. XXVI) and the Isle of Wight. If the Angles and Saxons could revisit the country they certainly would not recognise it; on the other hand, the Phoenicians might see little difference in the Cornish coast, except near St. Michael’s Mount. Even now there is good reason to suppose a slow sinking of south-eastern England. Since the end of the Ice Age there have been great changes in the levels of land and sea, and these have often had a more profound effect on the present appearance of the coast than have erosion and accretion.
BEACHES
If the beach along a fairly straight coast is examined, it will usually be noticed that if shingle is present it is at or near the top. Sand and finer particles occur lower down, and usually the fineness of material increases seawards. The cliffs behind the shingle may be of any kind, and are not necessarily the source of the shingle. After a severe storm much or all of the beach material can be removed, and the platform on which it rests exposed. In the succeeding normal weather, the beach will gradually accumulate again. Even after an ordinary blow the beach may be combed down, so that coarse and fine material are much more mixed.
When waves break, beach material, coarse and fine, is churned up. There is often some order and arrangement in this movement. If the waves are approaching the shore at right-angles, the pebbles and small stones move up and down the beach. The waves break and send up the beach sheets of water called the swash or send, which carry material upwards. Some of the water of the swash percolates into the beach, some returns to the sea as the backwash. This is nearly always less powerful than the swash, but in its deeper parts can move a good deal of material. If, however, the waves approach the beach obliquely, so also may they advance up it, and stones and sand are not merely carried upwards, but also sideways. When the swash dies out, the backwash returns directly down the slope, and any material moved by it travels in the same direction. Thus on open coasts on to which the waves come obliquely, there is a great deal of lateral displacement of beach material. This process is called beach-drifting, and is of the utmost importance. Its effect is often seen where groynes or breakwaters are built athwart the beach to hold material travelling along it by this process. The beach on one side of a groyne is usually higher than on the other, although often after a storm from a different quarter the high and low sides may temporarily change places.1
The waves also have a sorting effect, and drive the stones to higher parts of the beach. This process can often be seen in action while waves are breaking on a beach of mixed material. There is still another important factor. Around our coasts there is usually a noticeable difference in the level of the water at low and high tide. On the open coast the range seldom exceeds twenty feet, but in bays and gulfs it may be more. The highest rises and the lowest falls occur about the times of new and full moon. At the half-moons the difference between high and low water is small. Suppose it is now the moon’s first quarter, and the weather generally fair. The waves at both the morning and evening tide will reach to much about the same level. But in succeeding days the high waters will rise higher and the low waters will fall lower with each succeeding tide until the time of full moon.2 What effect will this have on the beach? If marked pebbles have been scattered near the water line at the time of the moon’s first quarter they will be seen not only to have moved along the beach if the waves are oblique, but to have been pushed up it by waves at each successive tide, and have gathered near the top, usually in an existing ridge. During the subsequent fall of tide-level after full moon, the pebbles are left stranded, and they may only just be reached again at the next period of springs at the time of new moon.
Thus, if nothing else happened, the pebbles might remain at the beach top for ever. But two other factors are likely to affect them. First, the swing of the tides from neaps to springs is only part of a larger swing that shows itself in particularly high tides near the equinoxes, and sometimes at other times of the year. Secondly, if a severe storm attacks a coast, especially at a period of big tides, shingle may be either swept far above its normal level or dragged in large quantities down the beach. An exceptional storm may overtop the highest beaches. The effect of ordinary storms is plain along any shingle beach, since the seaward face is frequently marked by minor ridges parallel to its length. These are either the heights reached by the last high tides, or the limits of recent storms.
Quite apart from these up-and-down and lateral movements of some beach material, vast quantities of finer stuff are moved alongshore by a different process. A bather on a sandy shelving beach in ordinary weather and in water three or four feet deep notices the lifting effect of the waves: the sand is at the same time somewhat disturbed about his feet. If he allows himself to float off such a beach, he notices that the tidal current carries him one way or another along it. The sand stirred up by the waves may, when a tidal current is running with any speed, also be carried sideways for a short distance. Picture this process during a tidal cycle, and during rough and stormy conditions, and it is at once apparent what vast quantities of fine material can be carried along a beach.
Current-action, however, only takes place under water and below the zone of wave-break. It operates on the higher parts of the beach only at or near high tide; on the lower parts, on an open coast the current may run one way at or near low water, and the opposite way at about high water. Two things at least follow—first, in the deeper water, material may move in different ways at different stages of the tide, and whether there is a balance of movement will depend upon the relative strengths of the flood and ebb currents. Secondly, on the parts of the beach covered only at high water, the movement of material is likely to be in one direction only—that of the current at the time of high water. The resultant process is called long-shore drifting. With beach-drifting it is of the utmost importance in the study of shoreline phenomena.
A wave breaks when it enters water the depth of which is approximately half its wave-length. Thus on a shallow coast, big waves break farther out than do small ones. When breaking offshore, the waves—just as on a beach—drive material up in front of them, so that sometimes ridges of sand and shingle are built some distance from the original shore. If a shingle ridge of this sort attains a fair degree of stability, it becomes an outer beach along which beach-drifting can take place. Hence the ridge may lengthen and become what is called an offshore bar (see here). If the process continues a lagoon-like expanse may be enclosed between it and the old shoreline. Offshore bars are seldom unbroken for long distances, since there are often gaps through which the tide enters and leaves the lagoon, in which marsh development is favoured.
If the supply of shingle is great, and if the lateral transport along a coast is marked, the shingle can accumulate in great forelands like Orfordness, Dungeness, the Crumbles, and the shingle ridges off the Culbin Sands and other parts of the Moray Firth shore. At an early stage a ridge is built. After a time the new shingle coming along the coast shallows the sea floor off the first ridge, so that the waves build another in front of it. This may go on until a whole series of such ridges is formed. It is often noticed that at one part of a shingle foreland the ridges run out to sea in such a way that it is clear they are suffering erosion, whereas at another part new shingle is accreting and being built up into ridges. A study of any big shingle foreland will illustrate this process, but there are few more striking examples than the shingle formation known as the Bar, near Nairn. Fig. 1 shows that it is composed of a number of individual ridges, the north-eastern ends of which are being eroded, whereas growth is continuous at the other end. In short, the whole structure is slowly shifting along the coast.
Along the south-west facing side of Dungeness there are many ridges running directly out to sea, and obviously at one time they continued for some distance. Erosion has cut them, and the material thus provided has travelled round the point of Dungeness and gradually helped to build the numerous ridges forming Denge Beach. Erosion is constantly taking place on the one side, accretion on the other.
FIG. 1.—The Bar (from Steers, Geogr. Journal, 1937).
The shingle that composes the banks and beaches comes partly from the erosion of cliffs, partly from boulder clay and other materials on the sea floor, and largely from glacial and gravel deposits, from which it has been swept by rivers in past times. Along our east and south coast it is mainly composed of flint which originally came from the Chalk. In west coast and Scottish beaches, the percentage of local rocks is far higher, and flint may be quite absent.
In its lateral travel alongshore, shingle often builds ridges or embankments, running across the mouths of rivers and inlets. Nearly all rivers are to some extent obstructed by a bar composed of shingle or sand, or both. When a bar is growing across a river mouth, the unattached end extends very much in the same manner as does a tip heap. At first the bar may be wholly below water; it gradually grows up to the surface. But whether above or below water, the free end tends to be turned inwards as a result of wave action. Many bars of this sort have on their landward side laterals or recurved ends (see here). Some bars grow forward, later turn inwards, and, after a time, grow forward again. It is easy to give general reasons for this—e.g. wave-attack in a storm—but it is extremely difficult to be precise. If the bar is obstructing a river, its form will depend in part on the power of the river to keep its mouth clear. Small streams like those at Chideock and other places on the Dorset coast are completely dammed. In others, e.g. the Exe and Teign, the river maintains a mouth; the Exe Bar is particularly interesting since it is double. At Orford Ness the shingle has not only formed a bar, but has grown into a great foreland and deflected the river for eleven miles. Some rivers like the Spey usually keep their mouths through shingle beaches in nearly the same place, whereas others, like the neighbouring Findhorn, by no means free from violent floods, are deflected.
Scolt Head Island and other features of the Norfolk coast and the Bar off Nairn on the Moray Firth are good examples of offshore bars, and, they, too, lengthen in the same way as ordinary beaches. Since they are offshore, they can send back long lateral ridges.
In bays and other inlets, shingle beaches are usually washed up at the head, to form what are called bay-head beaches. Normal beach-drifting for any distance along an indented coast is impossible. If, however, the bays are rather wider and more open, there is a certain amount of lateral travel of shingle in them, and it gathers at their leeward ends. The distribution of shingle along the several bays between Pwllheli and Penrhyndeudraeth is most instructive. Sometimes a ridge forms across the mid-part of a bay, such as Cemlyn Bay in Anglesey.
DUNES
Shingle ridges of various kinds have been discussed at some length because they form the “skeleton” combining the “flesh” of dunes and salt-marsh. By no means all dunes are built on shingle ridges, and many ridges have no dunes. But if a shingle ridge is being formed in a locality where an expanse of sand is bared at low tide, the wind blowing over the sand will carry much of it on to the ridge and deposit some of it in its interior and some on its surface. This may cause dune growth. But (see Chapter 7) the real dune-builders are the sea couch-grass (Agropyron junceiforme), marram-grass (Ammophila arenaria), and sea lyme-grass (Elymus arenarius) that take root in the shingle, send up shoots, and so begin to trap the sand. Marram-grass is remarkable in this way, thriving best where the sand supply is most prolific (Pl. III). If another shingle ridge is built in front of the old one, the same process will begin on that, and the dunes on the inner ridge will be, at any rate partially, deprived of their sand supply. The dunes may remain, but are often partly blown away, whereas those on the newer ridge increase. Under favourable circumstances they grow and become permanent features. In England dunes seldom exceed fifty or sixty feet in height if built up from sea-level.